We are interested in the evolution of plants and lichens. We use electron microscopy, DNA sequences and genomic data to resolve deep and shallow divergences in plants. Our recent interests are the functional genomics of cyanobacterial-plant symbioses, especially cycads (Zamia) and the hornwort Leiosporoceros dussii. We are developing genomic resources for hornworts, particularly the species Leiosporoceros dussii. We also interested in plant-microbial interaction in Canadian Arctic cryptogams